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I believe you need to disable the automatic hostnames.
Hope this helps!
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Disabling automatic hostnames has the impact, that I cannot use any Private Hosted Zone in AWS anymore. This is mentioned in same documentation you linked. ("If you use custom DNS domain names defined in a private hosted zone in Amazon Route 53, or [...], you must set both the enableDnsHostnames and enableDnsSupport attributes to true.")
However, your comment made me set this up, review all my config, and I found a typo in the FORWARD rules. FORWARD rules work now! A PHZ for PTR works too. So thanks a lot iBehr! Open is issue 2.